In the far eastern corner of the country of Yanato, it has been believed since ancient times that there are two worlds in this land: the world at the top where the gods live and the world at the foot of the mountain where humans live. Princess Sakuna, a high-ranking deity who lives in the world of the summit, was born to a war god and a fertility god but led a lazy life, wasting the grains her parents had stored. One day, due to some strange circumstances, she is banished from the city of the gods and sent to Hinoe Island, an isolated island inhabited by demons. In a barren land where there is barely enough food for the next day, she cultivates the soil, grows rice, and sets out to defeat the demons. The humans who have stumbled into the world of the gods and the novice fertility god begin their rice-husk-filled communal life together.